There is great geographical differentiation among the demes in the
frequency of the Standard chromosome arrangement (
). Since
is assumed to be equal to zero (no inbreeding
within demes), all of the departure from Hardy-Weinberg is a result of
among-deme differences.
Most of this geographical differentiation is a result of differences
among the four primary subdivisions (
). Relatively
minor differentiation is found among regions within a given
subdivision (
) and among demes within a region
(
).
Thus, an explanation for the chromosomal diversity that predicted great local differentiation and little or no differentiation at a large scale would be inconsistent with these observations.