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Thoreau coined the term ``succession'' as used in ecology to describe
the change in vegetation following natural disturbance. His first talk
on the subject, ``The succession of forest trees,'' was presented to
the Middlesex Agricultural Society, and it describes well the process
of old-field succession.
- The process starts with the conversion of tilled field to
pasture or its complete abandonment.
- White pine, pitch pine, and red cedar not only have small,
easily dispersed seeds, they are able to tolerate drought under full
sun, and are relatively unpalatable to grazing animals. Thus, pines
and red cedar tend to be the first trees that become established in
pastures. In recently tilled fields, however, red maple or birch
often establish more quickly and become dominant.
- Oak generally begins to appear in the understory once a
first-generation forest has been established.
Forest maps constructed by Charles Sprague Sargent and George Nichols
at the turn of the century identified much of New England as part of
the ``Northern Pine'' region (Figure 2), but
Extensive white pine forests were a transient phenomenon
generated by cultural and natural history ... For many stands of
pine and other pioneers, we can anticipate that with time and
natural development the vegetation will become something very
different ... The particular forest changes that have occurred
in the New England landscape over the last century were actually a
historical accident .... [6], p. 139.
And the effects of past land uses can linger for a long time. In spite
of more than a century of regrowth, forest soils in New England still
show a distinct plow horizon (Figure 3). Moreover,
the effects of past land use on vegetation are detectable more than a
century after it has been released from agricultural
use (Figure 4)
Figure 2:
One of Sargent's forest maps (from [9]).
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Figure 3:
Land use history and soil horizons in New England. The
horizon on the left shows the effects of plowing more than a century
after plowing stopped (from [4]).
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Figure 4:
Vegetation composition as a function of land use history at
two sites in southern New
England (from [4]).
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Next: Old growth forest in
Up: Thoreau's country: landscape change
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Kent Holsinger
2011-10-23